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Ph.D., University of Cambridge, England
Associate Professor
Department of Molecular Pharmacology, Physiology and Biotechnology
487 Biomedical Center
Tel: (401) 863-2574
Email: John_Marshall@Brown.edu
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In response to hormonal or synaptic stimulation, excitable cells (including
smooth muscle, cardiac muscle, and neurons) undergo a diversity of changes
in their electrical properties. Changes in channel activity are known to
regulate neuronal gene expression, cell death and communication. My lab is
studying the regulation of glutamate receptors and potassium channels by
protein kinases. Glutamate receptors play key roles in certain forms of synaptic
plasticity and the failure to regulate these receptors can cause seizure
disorders and neurodegenerative disease. We use patch-clamp techniques to
study the electrophysiological properties of cloned glutamate receptors heterologously
expressed in mammalian cell lines and Xenopus oocytes to determine how these
properties are modulated by kinases such as protein kinase A, CaM-Kinase
II and protein-tyrosine kinases (PTK). The PTK receptors, such as EGF, PDGF
and the neurotrophin receptors are expressed on fully differentiated neurons
and may also play an important role in regulating neuronal excitability.
In addition to the glutamate receptor studies we have recently cloned a novel
calcium-activated potassium channel. Site-directed mutagenesis methods are
being used to determine its structure and how the gating properties of this
channel are regulated by nitric oxide and other second messengers. Another
focus of the lab is the development of optical methods that utilize the cloned
jellyfish green fluorescent protein (GFP). We have engineered a chimeric
NMDA/ GFP receptor subunit that can be observed by fluorescence microscopy
while retaining its normal electrical properties. The tagging of receptors
with GFP will enable us to visualize the expression and localization of ion
channels in living cells.

Mehta S., Wu H., Garner CC, and Marshall J (2001) Intramolecular interactions
regulate SAP97 binding to kainate receptors. J. Biol. Chem. 2001 276: 16092-16099.
Liu, Y-F., Sudol, M., Marshall, J (2000) Assignment of molecular mechanisms
of neuronal death underlying Huntington's disease. J. Biol. Chem. 275, 19035-19040.
Blair, L.A.C., K.K. Bence and J. Marshall. (2000) Potentiation of neuronal
L calcium channels by IGF-1 requires phosphorylation of the a1 subunit on
a specific tyrosine residue. Neuron 27, 121-131.
Blair, L.A.C., K.K. Bence, T. Franke, D. Kaplan and J. Marshall. (1999)
Akt-dependent potentiation of L channels by IGF-1 is required for neuronal
survival. J. Neuroscience 19, 1940-51.
Garcia, E.P., Mehta, S., Blair, L.A.C., Wells, D.G., Shang, J., Fukushima.
T., Fallon, J.R., Garner, C.C., and Marshall, J. (1998) SAP90 binds and clusters
kainate receptors causing incomplete desensitization. Neuron 21, 727-39.
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